Trawling through decades of research and observation, he describes all sorts of behaviour in bonobos, chimpanzees, and monkeys (as well as non-primates such as dogs, elephants and dolphins) which clearly indicate that they have a well developed sense of right and wrong. Time and again, experiments have revealed that primates understand the value of co-operation in specific tasks; that they have a strong sense of fair play, an awareness of the permanence of death, and use their own experience and imagination in order to empathise with others.
MORE: http://www.independent.co.uk/arts-entertainment/books/reviews/review-the-bonobo-and-the-atheist-by-frans-de-waal-8581418.html
See also...
Where morals come from
http://lightyears.blogs.cnn.com/2013/04/12/science-seat-where-morals-come-from/?hpt=us_bn1